Universities in the USA Offering Full Scholarships- Application links, Requirements, How To Apply

Institution Program Program Link
Albert Einstein College of Medicine Biomedical Sciences https://www.einstein.yu.edu/education/phd/prospective-students/admissions.aspx
Auburn University Biological Sciences http://www.auburn.edu/cosam/departments/biology/grad/index.htm
Baylor College of Medicine Graduate School of Biomedical Sciences https://www.bcm.edu/education/schools/graduate-school-of-biomedical-sciences
Boston University Molecular Biology, Cell Biology, & Biochemistry http://www.bu.edu/mcbb/admissions/
Boston University Program in Biomedical Science http://www.bumc.bu.edu/gms/pibs/
Brandeis University Molecular and Cell Biology http://www.bio.brandeis.edu/grad/mcb/mcb_phd_rev.html
Brandeis University Neuroscience http://www.bio.brandeis.edu/grad/neuro/neuro_phd.html
Brown University Biomedical Engineering https://www.brown.edu/graduateprograms/biomedical-engineering-phd
Brown University Biotechnology https://www.brown.edu/graduateprograms/biomed-biotechnology-phd
Brown University Computational Biology https://www.brown.edu/graduateprograms/computational-biology-phd
Brown University Ecology and Evolutionary Biology https://www.brown.edu/graduateprograms/biomed-ecology-and-evolutionary-biology-phd
Brown University Molecular Biology, Cell Biology and Biochemistry https://www.brown.edu/graduateprograms/biomed-molecular-biology-cell-biology-and-biochemistry-phd
Brown University Molecular Pharmacology and Physiology https://www.brown.edu/graduateprograms/biomed-molecular-pharmacology-and-physiology-phd
Brown University Neuroscience https://www.brown.edu/graduateprograms/biomed-neuroscience-phd
Brown University Pathobiology https://www.brown.edu/graduateprograms/pathobiology-phd
Binghamton University – SUNY Behavioral Neuroscience https://www.binghamton.edu/psychology/graduate/behavioral-neuroscience/index.html
Binghamton University – SUNY Clinical Psychology https://www.binghamton.edu/psychology/graduate/clinical-psychology/index.html
Binghamton University – SUNY Biological Sciences https://www.binghamton.edu/biology/graduate/index.html
California State University Dominguez Hills Biology https://www.csudh.edu/gsr/graduate-studies/graduate-admission/#
California State University Sacramento Master’s of Stem Cel Biology https://www.csus.edu/college/natural-sciences-mathematics/biology/stem-cell-program.html
Caltech Biochemistry and Molecular Biophysics https://cce.caltech.edu/graduate/graduate-biochemistry-and-molecular-biophysics
Carnegie Mellon University Biological Sciences https://www.cmu.edu/bio/graduate/phd_program/admissions.html
Case Western Reserve University Biomedical Sciences Training Program https://case.edu/med/BSTP/apply.html
City University of New York Biology https://www.gc.cuny.edu/Prospective-Current-Students/Prospective-Students/Admissions#!
Clemson University Biological Sciences https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-biosci.html
Clemson University Environmental Toxicology https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-entox.html
Clemson University Microbiology https://www.clemson.edu/science/departments/biosci/academics/graduate/plan-study-micro.html
Cold Spring Harbor Laboratory PhD Program https://www.cshl.edu/education/phd-program/how-to-apply/
Colorado State University Biochemistry and Molecular Biology https://www.bmb.colostate.edu/
Columbia University Ecology, Evolution, and Environmental Biology http://e3b.columbia.edu/
Columbia University Medical School Biomedical Sciences Programs https://www.gsas.cuimc.columbia.edu/applying
Cornell University Neurobiology & Behavior https://nbb.cornell.edu/
Cornell University Biochemistry, Molecular, and Cell Biology https://bmcb.cornell.edu/
Cornell University Biomedical Engineering https://www.bme.cornell.edu/bme/programs/graduate-programs
Cornell University Entomology https://entomology.cals.cornell.edu/graduate/
Cornell University Genetics, Genomics, and Development https://gendev.cornell.edu/
Cornell University Plant Biology https://plantbio.cals.cornell.edu/academics/graduate/
Cornell University Plant Pathology https://pppmb.cals.cornell.edu/graduate/
Cornell University Biological and Biomedical sciences https://www.vet.cornell.edu/education/biomedical-biological-sciences-bbs-phd-program
Cornell University Ecology & Evolutionary Biology https://ecologyandevolution.cornell.edu/
Dartmouth Molecular & Cell Biology https://graduate.dartmouth.edu/mcb/
Dartmouth Program in Experimental & Molecular Medicine https://graduate.dartmouth.edu/pemm/
DePaul University Biological Sciences (MS) https://csh.depaul.edu/academics/biological-sciences/graduate/biological-sciences-ms/Pages/default.aspx
Drexel University Neuroscience (MS or PhD) https://drexel.edu/medicine/academics/graduate-school/neuroscience/
Drexel University Pharmacology and Physiology https://drexel.edu/medicine/academics/graduate-school/pharmacology-physiology/
Duke University Neurobiology https://www.neuro.duke.edu/
Duke University Biochemistry https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Cell & Molecular Biology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Developmental & Stem Cell Biology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Immunology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Molecular Cancer Biology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Molecular Genetics and Microbiology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Pharmacology https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Biology https://biology.duke.edu/graduate
Duke University Genetics and Genomics https://upg.duke.edu/
Eastern Virginia Medical School Biomedical Sciences https://www.evms.edu/education/doctoral_programs/biomedical_sciences/
Emory University Chemistry http://chemistry.emory.edu/home/graduate/apply.html
Emory University GDBBS – Biochemistry, Cell and Developmental Biology http://www.biomed.emory.edu/bcdb
Emory University GDBBS – Cancer Biology http://www.biomed.emory.edu/cb
Emory University GDBBS – Genetics and Molecular Biology http://www.biomed.emory.edu/gmb
Emory University GDBBS – Immunology and Molecular Pathogenesis http://www.biomed.emory.edu/imp
Emory University GDBBS – Microbiology and Molecular Genetics http://www.biomed.emory.edu/mmg
Emory University GDBBS – Molecular and Systems Pharmacology http://www.biomed.emory.edu/msp
Emory University GDBBS – Neuroscience http://www.biomed.emory.edu/ns
Emory University GDBBS – Population Biology, Ecology, and Evolution http://www.biomed.emory.edu/pbee
Florida State University Biological Sciences https://www.bio.fsu.edu/
Florida State University Biomedical Sciences https://med.fsu.edu/phd/home
Florida State University Neuroscience https://neuro.fsu.edu/gr/admission
Georgetown University Biomedical Graduate Education Pharmacology and Physiology https://pharmacology.georgetown.edu/program/
Georgetown University Biomedical Graduate Education Interdisciplinary Program in Neuroscience https://neuroscience.georgetown.edu/
Georgetown University Biomedical Graduate Education Tumor Biology https://tumorbiology.georgetown.edu/
Georgetown University Biomedical Graduate Education Biomedical masters degree programs https://biomedicalprograms.georgetown.edu/masters-programs/
Georgetown University Biology https://biology.georgetown.edu/graduate/
George Washington University IBS – Cancer Biology program https://smhs.gwu.edu/ibs/phd-programs-ibs/cancer-biology-phd-program
George Washington University IBS – Genomics & Bioinformatics https://smhs.gwu.edu/ibs/phd-programs-ibs/genomics-bioinformatics-phd-program
George Washington University IBS – Microbiology & Immunology https://smhs.gwu.edu/ibs/program/microbio-immunology
George Washington University IBS – Neuroscience https://smhs.gwu.edu/ibs/program/molecular-medicine/neurosciences
George Washington University IBS – Pharmacology & Physiology https://smhs.gwu.edu/ibs/program/molecular-medicine/pharma-physio
Gerstner Sloan Kettering Biomedical Sciences https://www.sloankettering.edu/gerstner/admissions/apply
Harvard University Biological and Biomedical Sciences http://www.hms.harvard.edu/dms/bbs/
Harvard University Biological Sciences in Dental Medicine https://gsas.harvard.edu/programs-of-study/all/biological-sciences-dental-medicine
Harvard University Biological Sciences in Public Health https://www.hsph.harvard.edu/biological-sciences/
Harvard University Biological Sciences in Public Health https://www.hsph.harvard.edu/biological-sciences/
Harvard University Biophysics https://gsas.harvard.edu/programs-of-study/all/biophysics
Harvard University Chemical Biology http://chembiophd.harvard.edu/
Harvard University Immunology http://dms.hms.harvard.edu/immunology/index.html
Harvard University Molecular and Cellular Biology https://gsas.harvard.edu/programs-of-study/all/molecular-and-cellular-biology
Harvard University Neuroscience http://dms.hms.harvard.edu/neuroscience/index.html
Harvard University Organismic & Evolutionary Biology https://gsas.harvard.edu/programs-of-study/all/organismic-and-evolutionary-biology
Harvard University Systems Biology http://sysbiophd.harvard.edu/prospective-students/admissions
Harvard University Virology http://dms.hms.harvard.edu/virology/index.html
Icahn School of Medicine at Mount Sinai Biomedical Science, Neuroscience http://icahn.mssm.edu/education/phd
Illinois State University Biological Sciences (PhD) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Neuroscience and Physiology) (PhD) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Molecular and Cellular Biology) (PhD) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Behavior,Ecology,Evolution and Systematics) (PhD) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Behavior,Ecology,Evolution and Systematics) (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Conservation Biology) (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Biomathematics) (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Bioenergy) (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Biotechnology) (MS) https://biology.illinoisstate.edu/graduate/
Illinois State University Biological Sciences (Neuroscience and Physiology) (MS) https://biology.illinoisstate.edu/graduate/
Indiana University Biology https://www.bio.indiana.edu/
Indiana University Cell, Molecular and Cancer Biology https://medicine.iu.edu/campuses/bloomington/graduate-programs/cell-molecular-and-cancer-biology-phd/
Indiana University Neuroscience https://neuroscience.indiana.edu/academics/how-to-apply/index.html
Indiana University School of Medicine Biomedical https://medicine.iu.edu/education/graduate-degrees/phd/admissions/
Iowa State University Ecology and Evolutionary Biology https://www.eeb.iastate.edu/admissions
Johns Hopkins School of Medicine Biochemistry, Cellular & Molecular Biology http://bcmb.bs.jhmi.edu/
Johns Hopkins School of Medicine Biological Chemistry http://biolchem.bs.jhmi.edu/Pages/application-information.aspx
Johns Hopkins School of Medicine Cellular and Molecular Medicine Program http://cmm.jhmi.edu/index.php/application-information/
Johns Hopkins School of Medicine Immunology https://gradimmunology.med.som.jhmi.edu/
Johns Hopkins School of Medicine Molecular Biophysics https://pmb.jhu.edu/admissions/
Johns Hopkins School of Medicine Neuroscience http://neuroscience.jhu.edu/graduate/apply
Johns Hopkins School of Medicine Pathobiology http://pathology.jhu.edu/department/training/pathbio_ProgramOverview.cfm
Johns Hopkins School of Medicine Pharmacology and Molecular Sciences http://www.hopkinsmedicine.org/pharmacology_molecular_sciences
Johns Hopkins Bloomberg School of Public Health Biochemistry and Molecular Biology http://www.jhu-bmb-phd.org/
Johns Hopkins University Biology (MS) https://bio.jhu.edu/undergraduate/masters-program/
Johns Hopkins University Cell, Molecular, Developmental Biology, & Biophysics https://cmdb.jhu.edu/
Johns Hopkins University Cellular and Molecular Physiology http://physiology.bs.jhmi.edu/academics/
Johns Hopkins University Human Genetics and Molecular Biology https://igm.jhmi.edu/humangenetics/welcome
Johns Hopkins University Molecular Biophysics http://pmcb.jhu.edu/
Kansas State University Biology https://www.k-state.edu/biology/
Massachusetts Institute of Technology Biological Engineering http://be.mit.edu/academic-programs/prospective-grad/graduate-application
Massachusetts Institute of Technology Biology https://biology.mit.edu/graduate/prospective-students/application-process/
Massachusetts Institute of Technology Brain and Cognitive Science https://bcs.mit.edu/academic-program/graduate/graduate-admissions
Mayo Clinic College of Medicine Biomedical Sciences https://college.mayo.edu/academics/biomedical-research-training/admissions-and-benefits/
Medical College of Wisconsin Interdisciplinary Program in Biomedical Sciences www.mcw.edu/idp
Medical College of Wisconsin Neuroscience www.mcw.edu/ndp
Medical College of Wisconsin Pharmacology & Toxicology www.mcw.edu/pharmtoxphd
Medical University of South Carolina Biomedical Sciences www.musc.edu/grad
Michigan Tech Biological Sciences http://www.mtu.edu/gradschool/programs/degrees/biological-sciences/
New York Medical College Basic Medical Sciences https://www.nymc.edu/graduate-school-of-basic-medical-sciences-gsbms/admissions–financial-aid/admission-requirements/
New York University Biology https://as.nyu.edu/biology/programs/phd.html
New York University Sackler Institute Biomedical Sciences https://med.nyu.edu/research/sackler-institute-graduate-biomedical-sciences/
NIH Oxford-Cambridge Scholars Program https://oxcam.gpp.nih.gov/
Northwestern University Neuroscience https://www.nuin.northwestern.edu/application-process-2/
Northwestern University Biological Sciences https://www.ibis.northwestern.edu/graduate/prospective/
Northwestern University Life Sciences http://www.feinberg.northwestern.edu/sites/dgp/prospective-students/phd-program/admissions.html
Ohio State University Biochemistry https://osbp.osu.edu/admissions
Ohio State University Evolution, Ecology and Organismal Biology https://eeob.osu.edu/grad/graduate-program
Ohio State University Microbiology https://microbiology.osu.edu/gradstudents/admissions
Ohio State University Molecular Genetics https://molgen.osu.edu/program-admissions-and-aid
Ohio State University Chemical and Biomolecular Engineering https://cbe.osu.edu/phd
Oregon State University Biochemistry and Biophysics https://biochem.oregonstate.edu/
Penn State College of Medicine Anatomy https://med.psu.edu/anatomy-phd
Penn State College of Medicine Biomedical Sciences https://med.psu.edu/biomedical-sciences-phd
Penn State College of Medicine Neuroscience https://med.psu.edu/neuroscience-phd
Penn State University Biochemistry, Microbiology, and Molecular Biology https://science.psu.edu/bmb
Penn State University Bioinformatics and Genomics https://www.huck.psu.edu/graduate-programs/bioinformatics-and-genomics/
Penn State University Biology https://science.psu.edu/bio
Princeton University Ecology and Evolutionary Biology https://eeb.princeton.edu/
Princeton University Molecular Biology https://molbio.princeton.edu/
Princeton University Neuroscience http://pni.princeton.edu/
Purdue University Biological Sciences https://www.bio.purdue.edu/
Purdue University Chemistry https://www.chem.purdue.edu/faq/index.php#gre
Richard Gilder Graduate School- American Museum of Natural History Comparative Biology https://www.amnh.org/research/richard-gilder-graduate-school/apply
Rockefeller University Graduate Program https://graduateapplication.rockefeller.edu/
Roswell Park Cancer Institute Cancer Sciences https://www.roswellpark.org/education/phd-program
Rutgers University Biochemistry (Molecular Biosciences) https://molbiosci.rutgers.edu/
Rutgers University Cell and Developmental Biology (Molecular Biosciences) https://molbiosci.rutgers.edu/
Rutgers University Microbiology & Molecular Genetics (Molecular Biosciences) https://molbiosci.rutgers.edu/
Rutgers University Neuroscience http://rwjms.rutgers.edu/gsbs/programs/neuroscience/index.html
Rutgers University Pharmacology, Cellular & Molecular (Molecular Biosciences) https://molbiosci.rutgers.edu/
Rutgers University Physiology & Integrative Biology (Molecular Biosciences) https://molbiosci.rutgers.edu/
Rutgers University Toxicology https://eohsi.rutgers.edu/divisions/toxicology/
Rutgers University (Newark) Cell Biology, Neuroscience and Physiology (Newark) http://njms.rutgers.edu/sgs/prospective_students/multidisciplinary_cbnp2.php
Rutgers University (Newark) Immunity and Inflammation (Newark) http://njms.rutgers.edu/sgs/current_students/multidisciplinary_i3.php
Rutgers University (Newark) Molecular Biology Genetics and Cancer (Newark) http://njms.rutgers.edu/sgs/prospective_students/multidisciplinary_mbgc.php
San Francisco State University Biology (Master’s) http://biology.sfsu.edu/content/applying-sfsu-biology-graduate-programs
Scripps Research Institute Skaggs Graduate School https://education.scripps.edu/graduate/admissions/faqs/
Stanford University Bioengineering https://bioengineering.stanford.edu/
Stanford University Biosciences https://biosciences.stanford.edu/prospective-students/how-to-apply/
Stanford University Chemistry https://chemistry.stanford.edu/academics/phd-program/admissions/how-apply
Stowers Institute for Medical Research Graduate Program http://www.stowers.org/gradschool/admissions
Texas A&M University Biology https://www.bio.tamu.edu/
Texas A&M University Ecology and Evolutionary Biology https://eeb.tamu.edu/
Texas A&M University Genetics https://genetics.tamu.edu/
Thomas Jefferson University Graduate School of Biomedical Sciences https://www.jefferson.edu/university/life-sciences.html
Tufts University Conservation Medicine (MS) https://grad.vet.tufts.edu/ms-conservation-medicine/
University of Alabama Birmingham Biomedical Sciences http://www.uab.edu/gbs/home/admissions
University of Arizona Biological and Biomedical Sciences https://abbs.arizona.edu/
University of Arizona School of Plant Sciences https://cals.arizona.edu/spls/home
University of Buffalo Program in Biomedical Science (PPBS) http://medicine.buffalo.edu/phdprogram.html
University of Buffalo Microbiology and Immunology http://medicine.buffalo.edu/departments/micro/education/phd.html
University of California Berkeley Biophysics https://biophysics.berkeley.edu/admissions/
University of California Berkeley Environmental Science, Policy, and Management https://ourenvironment.berkeley.edu/graduate-programs/faq
University of California Berkeley Integrative Biology (IB) http://ib.berkeley.edu/
University of California Berkeley Microbiology http://plantandmicrobiology.berkeley.edu/graduate-program-microbiology
University of California Berkeley Molecular and Cell Biology https://mcb.berkeley.edu/grad/prospective-graduates/admissions-and-recruitment
University of California Berkeley Neuroscience https://neuroscience.berkeley.edu/ph-d-program/
University of California Berkeley Plant Biology http://plantandmicrobiology.berkeley.edu/plant-biology-0
University of California Berkeley Bioengineering (joint with UC San Francisco) https://bioegrad.berkeley.edu/prospectivegrads/admissions
University of California Davis Biochemistry, Molecular, Cellular and Developmental Biology http://bmcdb.ucdavis.edu/admissions/index.html
University of California Davis Ecology https://ecology.ucdavis.edu/
University of California Davis Medical Microbiology & Immunology https://health.ucdavis.edu/medmicro/
University of California Davis Molecular, Cellular, and Integrative Physiology https://mcip.ucdavis.edu/
University of California Davis Nutritional Biology https://ggnb.ucdavis.edu/
University of California Davis Plant Biology https://pbi.ucdavis.edu/about
University of California Davis Plant Pathology http://plantpathology.ucdavis.edu/gradindex.cfm
University of California Davis Population Biology https://cpb.ucdavis.edu/
University of California Davis Horticulture and Agronomy https://ggha.ucdavis.edu/
University of California Davis Integrative Pathobiology https://integrativepath.vetmed.ucdavis.edu/
University of California Davis Integrative Genetics and Genomics https://igg.ucdavis.edu/
University of California Davis Population Biology https://pbg.ucdavis.edu/
University of California Davis Neuroscience https://grad.neuroscience.ucdavis.edu/apply
University of California Irvine Ecology and Evolutionary Biology https://ecoevo.bio.uci.edu/graduates/admissions/
University of California Los Angeles Biochemistry, Molecular, and Structural Biology https://bmsb.chem.ucla.edu/how-apply
University of California Los Angeles Bioscience (Varies by program) http://bioscience.ucla.edu/application-procedure
University of California Los Angeles Ecology and Evoltionary Biology https://www.eeb.ucla.edu/graduate/
University of California Merced Quantitative and Systems Biology http://qsb.ucmerced.edu/
University of California San Diego Biomedical Sciences http://biomedsci.ucsd.edu/how-to-apply/index.html
University of California San Francisco Biomedical Sciences https://bms.ucsf.edu/
University of California San Francisco Chemistry and Chemical Biology https://ccb.ucsf.edu/admissions
University of California San Francisco Developmental and Stem Cell Biology https://dscb.ucsf.edu/
University of California San Francisco Integrative Program in Quantitative Biology https://ipqb.ucsf.edu/
University of California San Francisco Neuroscience http://neurograd.ucsf.edu/about-neuroscience-graduate-program
University of California San Francisco Oral and Craniofacial Sciences https://dentistry.ucsf.edu/programs/oral-cranio-phd
University of California San Francisco Pharmaceutical Sciences and Pharmacogenomics https://pspg.ucsf.edu/
University of California San Francisco Rehabilitation Science https://ptrehab.ucsf.edu/rehabilitation-science-phd
University of California San Francisco Tetrad https://tetrad.ucsf.edu/admissions
University of California Santa Barbara Biomolecular Science and Engineering https://www.bmse.ucsb.edu/
University of California Santa Barbara Molecular, Cellular, and Developmental Biology https://www.mcdb.ucsb.edu/academics/graduate
University of California Santa Cruz Biomolecular Engineering and Bioinformatics https://pbse.ucsc.edu/index.html
University of California Santa Cruz Chemical Biology, Biochemistry and Biophysics https://pbse.ucsc.edu/index.html
University of California Santa Cruz Microbiology and Environmental Toxicology https://pbse.ucsc.edu/index.html
University of California Santa Cruz Molecular, Cell, and Developmental Biology https://pbse.ucsc.edu/index.html
University of Chicago Biosciences Programs https://biosciences.uchicago.edu/programs
University of Chicago Ecology and Evolution https://pondside.uchicago.edu/ee/index.html
University of Colorado Anschutz Medical Campus Biomedical Sciences https://www.ucdenver.edu/academics/colleges/Graduate-School/academic-programs/Biomedical/Pages/home.aspx
University of Colorado Anschutz Medical Campus Cancer Biology https://www.ucdenver.edu/academics/colleges/Graduate-School/academic-programs/cancerbiology/Pages/home.aspx
University of Colorado Anschutz Medical Campus Cell Biology, Stem Cells and Development https://www.ucdenver.edu/academics/colleges/medicalschool/programs/CSD/Pages/home.aspx
University of Colorado Anschutz Medical Campus Immunology https://medschool.cuanschutz.edu/immunology-and-microbiology
University of Colorado Anschutz Medical Campus Microbiology https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Molecular Biology https://www.ucdenver.edu/academics/colleges/medicalschool/programs/Molbio/Pages/home.aspx
University of Colorado Anschutz Medical Campus Pharmaceutical Sciences https://www.ucdenver.edu/academics/colleges/pharmacy/AcademicPrograms/PhDPrograms/PhDPharmaceuticalSciences/Pages/PhDPharmaceuticalSciences.aspx
University of Colorado Anschutz Medical Campus Pharmacology https://medschool.cuanschutz.edu/pharmacology
University of Colorado Anschutz Medical Campus Toxicology https://www.ucdenver.edu/ACADEMICS/COLLEGES/PHARMACY/ACADEMICPROGRAMS/PHDPROGRAMS/PHDTOXICOLOGY/Pages/PhDToxicology.aspx
University of Colorado at Boulder Molecular, Cellular & Developmental Biology https://www.colorado.edu/mcdb/graduate/application-details-info
University of Colorado at Boulder Ecology and Evolutionary Biology https://www.colorado.edu/ebio/
University of Connecticut Molecular and Cell Biology https://mcb.uconn.edu/application-information-mcb-ph-d-and-ms-graduate-programs-2/
University of Denver Cell and Molecular Biology (MS & PhD) https://www.du.edu/academics/admission-steps/biological-sciences-cell-and-molecular-biology-phd-2021-2022
University of Denver Ecology and Evolutionary Biology (MS & PhD) https://www.du.edu/academics/admission-steps/biological-sciences-biology-ecology-and-evolution-phd-2021-2022
University of Georgia Cellular Biology (MS) https://cellbio.uga.edu/graduate-program
University of Georgia Integrated Life Sciences https://ils.uga.edu/
University of Illinois at Chicago Pharmaceutical Sciences Program https://pharmacy.uic.edu/programs/graduate-programs/phd-programs/phd-in-pharmaceutical-sciences/
University of Illinois at Urbana-Champaign Chemistry http://chemistry.illinois.edu/admissions/graduate-admissions
University of Illinois at Urbana-Champaign Molecular and Cellular Biology https://mcb.illinois.edu/graduate/gradprospect/
University of Iowa Biomedical Science https://medicine.uiowa.edu/biomed
University of Iowa Biomedical Engineering Program https://bme.engineering.uiowa.edu/graduate/bme-graduate-program-phd
University of Iowa Neuroscience https://neuroscience.grad.uiowa.edu/
University of Kansas Ecology and Evolutionary Biology https://eeb.ku.edu/how-apply
University of Kansas Molecular Biosciences https://molecularbiosciences.ku.edu/
University of Kansas Bioengineering http://bio.engr.ku.edu/
University of Kansas Medical Center Interdisciplinary Graduate Program in Biomedical Sciences http://www.kumc.edu/igpbs.html
University of Kentucky Biology https://bio.as.uky.edu/grad-program
University of Louisville Biomedical Sciences-IPIBS http://louisville.edu/medicine/ipibs/for-students/application-information/application-process
University of Maine School of Biology and Ecology https://sbe.umaine.edu/
University of Maryland Baltimore Varies by program (see link) https://lifesciences.umaryland.edu/prospective/Admissions/
University of Maryland Baltimore Chemical Engineering https://cbee.umbc.edu/academics/graduate-application/
University of Massachusetts Amherst Environmental Conservation https://eco.umass.edu/
University of Massachusetts Amherst Molecular and Cellular Biology http://gpls.cns.umass.edu/mcb
University of Massachusetts Amherst Neuroscience and Behavior http://gpls.cns.umass.edu/nsb
University of Massachusetts Amherst Organismic & Evolutionary Biology https://gpls.cns.umass.edu/oeb
University of Massachusetts Amherst Plant Biology http://gpls.cns.umass.edu/pb
University of Massachusetts Medical School Biomedical Sciences https://www.umassmed.edu/gsbs/
University of Memphis Biological Sciences https://www.memphis.edu/biology/graduate/
University of Miami Biology https://biology.as.miami.edu/graduate/index.html
University of Michigan Biophysics https://lsa.umich.edu/biophysics/graduates/admissions.html
University of Michigan Chemical Biology http://www.chembio.umich.edu/
University of Michigan Ecology and Evolutionary Biology https://lsa.umich.edu/eeb
University of Michigan Program in Biomedical Science https://medicine.umich.edu/medschool/education/phd-programs/phd-admissions
University of Minnesota Ecology and Evolutionary Biology https://cbs.umn.edu/academics/departments/eeb/graduate/graduate-admissions
University of Minnesota Biochemistry, Molecular Biology & Biophysics https://www.med.umn.edu/education-training/graduate-programs
University of Minnesota Integrative Biology and Physiology https://www.physiology.umn.edu/degrees-and-programs/graduate-program
University of Minnesota Microbiology, Immunology and Cancer Biology http://www.micab.umn.edu/
University of Minnesota Molecular, Cellular, Developmental Biology & Genetics https://www.med.umn.edu/education-training/graduate-programs
University of Minnesota Neuroscience https://med.umn.edu/neuroscience
University of Minnesota Pharmacology https://www.med.umn.edu/education-training/graduate-programs
University of Minnesota Plant and Microbial Biology https://cbs.umn.edu/academics/departments/pmb/graduate-education/admissions
University of Missouri Division of Biological Sciences http://biology.missouri.edu/graduate-studies/apply/
University of Montana Organismal Biology, Ecology, and Evolution http://hs.umt.edu/dbs/grad-programs/OBE/default.php
University of Nebraska Medical Center Interdisciplinary Graduate Program in Biomedical Sciences https://www.unmc.edu/gradstudies/programs/phd/interdisciplinary-biomedical-sciences.html
University of Nevada, Reno’s Integrative Neuroscience Program https://www.unr.edu/neuroscience/graduate-program
University of New Mexico Biology http://biology.unm.edu/graduate/graduate-admissions.shtml
University of North Carolina at Chapel Hill Biological and Biomedical Sciences Program https://bbsp.unc.edu/
University of North Carolina at Chapel Hill Biology (incl QBio and EEB) https://bio.unc.edu/graduate/
University of Notre damme Biological Sciences https://biology.nd.edu/faculty-research/areas-of-research/
University of Oklahoma Microbiology (MS and PhD) http://www.ou.edu/admissions/academics/arts-sciences/microbiology
University of Oklahoma Microbiology and Plant Biology (MS and PhD) ou.edu/cas/mpbio
University of Oklahoma Health Sciences Graduate Program in Biomedical Sciences http://graduate.ouhsc.edu/GraduatePrograms/PhDPrograms/GraduatePrograminBiomedicalSciences.aspx
University of Oregon Biology https://biology.uoregon.edu/graduate-studies/apply/
University of Oregon Chemistry and Biochemistry https://chemistry.uoregon.edu/
University of Pennsylvania Biology https://www.bio.upenn.edu/graduate/
University of Pennsylvania Biomedical Graduate Studies http://www.med.upenn.edu/bgs/
University of Pittsburgh Biological Sciences http://www.biology.pitt.edu/graduate/how-apply
University of Pittsburgh Biomedical Informatics https://www.dbmi.pitt.edu/content/doctoral-degree-0
University of Pittsburgh Cell Biology and Molecular Physiology http://www.gradbiomed.pitt.edu/programs/cell-biology-and-molecular-physiology
University of Pittsburgh Cellular and Molecular Pathology http://www.gradbiomed.pitt.edu/programs/cellular-molecular-pathology
University of Pittsburgh Clinical and Translational Science https://www.icre.pitt.edu/ClinTransPhD/index.html
University of Pittsburgh Computational Biology http://www.compbio.cmu.edu/
University of Pittsburgh Integrative Systems Biology https://www.isb.pitt.edu/
University of Pittsburgh Microbiology and Immunology http://pmi.pitt.edu/
University of Pittsburgh Molecular Biophysics & Structural Biology http://www.mbsb.pitt.edu/
University of Pittsburgh Molecular Genetics & Developmental Biology http://www.gradbiomed.pitt.edu/programs/molecular-genetics-development-biology
University of Pittsburgh Molecular Pharmacology http://www.gradbiomed.pitt.edu/programs/molecular-pharmacology
University of Pittsburgh Neurobiology https://www.cnup.pitt.edu/html/CNUP_Homepage.html
University of Rochester Medical Center Biochemistry & Molecular Biology https://www.urmc.rochester.edu/education/graduate/phd/biochemistry.aspx
University of Rochester Medical Center Biophysics, Structural & Computational Biology https://www.urmc.rochester.edu/education/graduate/phd/biophysics.aspx
University of Rochester Medical Center Cell Biology of Disease https://www.urmc.rochester.edu/education/graduate/phd/pathology.aspx
University of Rochester Medical Center Cellular and Molecular Pharmacology and Physiology https://www.urmc.rochester.edu/education/graduate/phd/pharmacology-and-physiology.aspx
University of Rochester Medical Center Genetics, Development and Stem Cells https://www.urmc.rochester.edu/education/graduate/phd/genetics.aspx
University of Rochester Medical Center Immunology, Microbiology & Virology https://www.urmc.rochester.edu/education/graduate/phd/immunology-microbiology-virology.aspx
University of Rochester Medical Center Neuroscience https://www.urmc.rochester.edu/education/graduate/phd/neurosciences.aspx
University of Rochester Medical Center Toxicology https://www.urmc.rochester.edu/education/graduate/phd/toxicology.aspx
University of Rochester Medical Center Translational Biomedical Science https://www.urmc.rochester.edu/education/graduate/phd/translational-biomedical-science.aspx
University of South Dakota School of Medicine Basic Biomedical Sciences https://www.usd.edu/medicine/basic-biomedical-sciences
University of Southern California Biomedical and Biological Sciences https://keck.usc.edu/pibbs/
University of Southern California Pharmaceutical and Translational Sciences https://pharmgradprograms.usc.edu/phts/phd/admission/apply/
University of South Carolina Biological Sciences https://www.sc.edu/study/colleges_schools/graduate_school/apply/degree_programs-application-requirements/biological-sciences-phd/index.php
University of Tennessee Ecology and Evolutionary Biology https://eeb.utk.edu/
University of Tennessee Microbiology https://micro.utk.edu/
UT Health San Antonio Integrated Biomedical Sciences (IBMS) Program https://www.uthscsa.edu/academics/biomedical-sciences/programs/integrated
UT Health San Antonio Translational Science Program https://www.uthscsa.edu/academics/biomedical-sciences/programs/translational-science-phd
University of Texas MD Anderson Cancer Center UTHealth Graduate School of Biomedical Sciences https://gsbs.uth.edu/
University of Texas Southwestern Medical School Graduate School of Biomedical Sciences (Umbrella PhD and MS) http://www.utsouthwestern.edu/education/graduate-school/application-and-admissions/
University of Utah Bioscience Program (Umbrella PhD) https://bioscience.utah.edu/#:~:text=The%20Bioscience%20PhD%20Programs%20are,at%20the%20University%20of%20Utah.
University of Utah Molecular, Cellular, and Evolutionary Biology https://www.biology.utah.edu/graduate/mceb/apply.php
University of Utah Ecology, Evolution, and Organismal Biology https://www.biology.utah.edu/graduate/eeob/apply.php
University of Vermont Biology https://www.uvm.edu/cas/biology/application-finances-support
University of Vermont Cellular, Molecular, and Biomedical Sciences https://www.uvm.edu/cmb
University of Vermont Neuroscience https://www.uvm.edu/neurosciencegrad
University of Virginia Biomedical Sciences http://bims.virginia.edu/
University of Virginia Biology http://bio.as.virginia.edu
University of Virginia Ecology and Evolutionary Biology https://www.eebvirginia.org/
University of Virginia Psychology https://psychology.as.virginia.edu/graduate-studies
University of Virginia Biomedical Engineering https://engineering.virginia.edu/departments/biomedical-engineering
University of Washington Neuroscience https://depts.washington.edu/neurogrd/prospective-students/admissions/
University of Washington Biochemistry https://depts.washington.edu/biowww/
University of Washington Biology https://www.biology.washington.edu/
University of Washington Chemical Engineering https://www.cheme.washington.edu/
University of Washington Genome Sciences https://www.gs.washington.edu/academics/gradprogram/index.htm
University of Washington Immunology http://www.immunology.washington.edu/graduate-program/admissions/admissions-faqs/
University of Washington Molecular Medicine and Mechanisms of Disease http://www.pathology.washington.edu/academics/mmtp/phd-program/
University of Washington Molecular and Cellular Biology https://www.washington.edu/students/gencat/academic/mcb.html
University of Wisconsin-Madison Cell and Molecular Biology https://cmb.wisc.edu/admissions/
University of Wisconsin-Madison Endocrinology https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
University of Wisconsin-Madison Integrated Program in Biochemistry https://ipib.wisc.edu/
University of Wisconsin-Madison Microbiology https://microbiology.wisc.edu/ps_apply.php
University of Wisconsin-Madison Neuroscience https://ntp.neuroscience.wisc.edu/
University of Wisconsin-Madison Nutritional Sciences https://nutrisci.wisc.edu/graduate/
University of Wisconsin-Madison Pharmacology https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
Van Andel Institute Molecular and Cellular Biology https://vaigs.vai.org/admissions/apply/
Vanderbilt University Biological Sciences https://as.vanderbilt.edu/biosci/graduate/admission.php
Vanderbilt University Interdisciplinary Graduate Program https://medschool.vanderbilt.edu/igp/
Vanderbilt University Quantitative and Chemical Biology https://medschool.vanderbilt.edu/qcb/
Vanderbilt University Neuroscience https://medschool.vanderbilt.edu/brain-institute/new-vbi-homepage-inprogress/join-our-graduate-program/
Virginia Commonwealth University Biomedical Sciences Doctoral Portal https://medschool.vcu.edu/education/bsdp/
Virginia Commonwealth University Clinical and Translational Research, Cancer and Molecular Medicine http://bulletin.vcu.edu/graduate/office-research/center-clinical-translational-research/clinical-translational-sciences-phd-concentration-cancer-molecular-medicine/
Virginia Tech Biochemistry https://www.biochem.vt.edu/
Virginia Tech Biological Sciences https://www.biol.vt.edu/
Wake Forest School of Medicine Biomedical Graduate Programs (multiple programs) https://school.wakehealth.edu/Education-and-Training/Graduate-Programs
Wake Forest School of Medicine Molecular and Cellular Biosciences Track https://school.wakehealth.edu/Education-and-Training/Graduate-Programs/Molecular-and-Cellular-Biosciences-Track#:~:text=The%20Molecular%20and%20Cellular%20Bioscience,to%20a%20specific%20research%20topic.
Washington State University Biological Sciences https://sbs.wsu.edu/
Washington State University Biological Systems Engineering https://bsyse.wsu.edu/graduate-program/how-to-apply/
Washington University in St. Louis DBBS – Biochemistry, Biophysics & Structural Biology http://dbbs.wustl.edu/divprograms/biophysics/Pages/BBSB.aspx
Washington University in St. Louis DBBS – Developmental, Regenerative & Stem Cell Biology http://www.dbbs.wustl.edu/divprograms/devbio/Pages/default.aspx
Washington University in St. Louis DBBS – Cancer Biology http://dbbs.wustl.edu/divprograms/cancerbiology/Pages/default.aspx
Washington University in St. Louis DBBS – Computational & Systems Biology http://dbbs.wustl.edu/divprograms/compbio/Pages/default.aspx
Washington University in St. Louis DBBS – Evolution, Ecology & Population Biology http://www.dbbs.wustl.edu/divprograms/eepb/Pages/default.aspx
Washington University in St. Louis DBBS – Human & Statistical Genetics http://dbbs.wustl.edu/divprograms/hsg/Pages/default.aspx
Washington University in St. Louis DBBS – Immunology http://dbbs.wustl.edu/divprograms/immunology/Pages/default.aspx
Washington University in St. Louis DBBS – Molecular Genetics & Genomics http://www.dbbs.wustl.edu/divprograms/genetics/Pages/default.aspx
Washington University in St. Louis DBBS – Molecular Microbiology & Microbial Pathogenesis http://www.dbbs.wustl.edu/divprograms/micro/Pages/default.aspx
Washington University in St. Louis DBBS – Neurosciences http://dbbs.wustl.edu/divprograms/neuro/Pages/default.aspx
Washington University in St. Louis DBBS – Plant & Microbial Biosciences http://dbbs.wustl.edu/divprograms/PlantMicroBioSci/Pages/default.aspx
Washington University in St. Louis DDBS – Molecular Cell Biology http://www.dbbs.wustl.edu/divprograms/cellbio/Pages/default.aspx
Wayne State University Biochemistry, Immunology and Microbiology https://biochemmicroimmuno.med.wayne.edu/
Washington University in St. Louis DDBS – Biomedical Informatics & Data Science http://dbbs.wustl.edu/divprograms/BIDS/Pages/default.aspx
Weill Cornell Graduate School of Medical Sciences BCMB Allied Program https://gradschool.weill.cornell.edu/bcmb-allied
Weill Cornell Graduate School of Medical Sciences Immunology & Microbial Pathogenesis Program https://gradschool.weill.cornell.edu/programs/immunology-microbial-pathogenesis
Weill Cornell Graduate School of Medical Sciences Neuroscience Program https://gradschool.weill.cornell.edu/programs/neuroscience
Weill Cornell Graduate School of Medical Sciences Pharmacology Program https://gradschool.weill.cornell.edu/programs/pharmacology
Weill Cornell Graduate School of Medical Sciences Physiology, Biophysics and Systems Biology Program https://gradschool.weill.cornell.edu/programs/physiology-biophysics-systems-biology
West Virginia University Biomedical Sciences https://www.hsc.wvu.edu/resoff/graduate-education/phd-programs/biomedical-sciences/
Wright State University Biological Sciences (MS) https://science-math.wright.edu/biology/master-of-science-in-biological-sciences
Yale University Combined Program in Biological & Biomedical Sciences https://medicine.yale.edu/bbs/
Updated February 13, 2021
Please email Joshua Hall ([email protected]) or tweet to @jdhallphd for additions or corrections to this list.

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